Lesson Notes By Weeks and Term v5 - Grade 11

Transport systems in plants – Week 1 focus

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Subject: Life Sciences

Class: Grade 11

Term: 2nd Term

Week: 1

Theme: General lesson support

Lesson Video

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Performance objectives

Lesson summary

This week, we begin exploring how plants, the foundation of almost all ecosystems, manage to transport essential substances throughout their bodies. Just like our own bodies need a circulatory system to deliver nutrients and remove waste, plants have their own intricate transport systems. Understanding these systems is crucial because it helps us appreciate how plants grow, survive, and contribute to our environment. From the towering baobab trees of Limpopo to the mielies in a Free State farm, the principles of plant transport are at play.

Lesson notes

2.1 Vascular Tissues: Xylem and Phloem Plants, unlike simple organisms, have specialized tissues for long-distance transport: xylem and phloem. These are collectively known as vascular tissues.

Xylem: The xylem is primarily responsible for transporting water and dissolved mineral salts (nutrients) from the roots to the rest of the plant, especially the leaves where photosynthesis occurs. The xylem is composed of dead cells called tracheids and vessel elements in flowering plants. These cells are elongated and hollow, forming continuous tubes that allow for efficient water flow. The walls of xylem cells are reinforced with lignin, providing structural support to the plant. Lignin also makes the xylem walls impermeable to water, preventing leakage. Think of the xylem as the plant's "water pipes." Phloem: The phloem transports sugars (produced during photosynthesis) and other organic compounds (e.g., amino acids, hormones) from the leaves (source) to other parts of the plant where they are needed for growth, storage, or other metabolic processes (sinks). This process is known as translocation. The phloem consists of living cells called sieve tube elements and companion cells. Sieve tube elements are connected end-to-end by sieve plates, which have pores that allow the flow of sap. Companion cells are closely associated with sieve tube elements and provide them with metabolic support. Unlike xylem, phloem cells are alive. Imagine the phloem as the plant's "food delivery system." 2.2 Water Absorption by Roots Plants absorb water primarily through their root hairs, which are tiny, hair-like extensions of epidermal cells located near the tips of roots. These root hairs greatly increase the surface area available for water absorption. The movement of water into root hair cells is driven by osmosis. Osmosis is the movement of water from a region of high water potential (low solute concentration) to a region of low water potential (high solute concentration) across a semi-permeable membrane. The soil surrounding the roots typically has a higher water potential than the cytoplasm of the root hair cells. This difference in water potential is due to the presence of dissolved mineral salts and sugars within the root cells. As a result, water moves from the soil into the root hair cells by osmosis.

Example: Consider a maize plant growing in a field. The soil is well-watered, and the concentration of solutes (e.g., fertilizers, salts) is relatively low. The cytoplasm of the root hair cells contains dissolved sugars and mineral salts, resulting in a lower water potential. Water will therefore move from the soil into the root hair cells, following the water potential gradient. 2.3 Pathways of Water Movement: Apoplast and Symplast Once water enters the root hair cells, it needs to move across the root cortex to reach the xylem. There are two main pathways for water movement: Apoplast Pathway: This pathway involves the movement of water through the cell walls and intercellular spaces of the root cortex cells. The apoplast pathway is relatively fast because water does not need to cross any cell membranes.

However, the apoplast pathway is blocked at the Casparian strip, a band of waterproof material (suberin) located in the cell walls of the endodermis (the innermost layer of the cortex). The Casparian strip forces water to enter the symplast pathway.

Symplast Pathway: This pathway involves the movement of water from cell to cell through the cytoplasm, via plasmodesmata (small channels that connect the cytoplasm of adjacent cells). The symplast pathway is slower than the apoplast pathway because water needs to cross cell membranes.

However, it allows the plant to control which substances enter the xylem, preventing harmful substances from reaching the leaves.

Example: Think of the Casparian strip like a security checkpoint. Water traveling through the apoplast (like walking through a public area) is stopped by the Casparian strip and forced to enter the symplast (like going through security and entering a restricted area). This allows the plant to "check" what is dissolved in the water before it enters the xylem. 2.4 Water Transport Through the Xylem Water is transported through the xylem from the roots to the leaves against gravity. This is primarily driven by the transpiration-cohesion-tension mechanism.

Transpiration: Transpiration is the loss of water vapor from the leaves through small pores called stomata. As water evaporates from the leaves, it creates a negative pressure (tension) in the xylem.

Cohesion: Water molecules are attracted to each other due to cohesion, which is the attraction between like molecules (in this case, water molecules). This cohesion creates a continuous column of water in the xylem from the roots to the leaves.

Adhesion: Water molecules are also attracted to the walls of the xylem vessels due to adhesion, which is the attraction between unlike molecules (in this case, water and the xylem walls).